When the condenser mic is off, R1 provides about 186 uA of current into the base of Q1. If Q1 has a beta (current gain) of at least 50, then Q1 will be close to saturation, meaning that it's collector voltage (which is TRI input of the '555) is close to 0 V. In this condition, the base voltage of Q1 is around +0.7 V.
Updated this section
OP did not provide any information on the microphone, so I'm going to have to make some assumptions. When a sound is detected by the mic, it generates an AC waveform whose frequency is that of the incoming sound wave.
This should alternately pull the base to close to 0 V assuming the mic can sink the 186 uA from R1, and then let it rise to 0.7 V . This actions turns Q1 off and on at the sound frequency, which subsequently causes the collector (TRI input of the '555) to swing between 9 V and 0 V, which triggers the timer.
All this said, it looks like the sense of the TRI input is backwards from what it should be. I also think there should be some filtering - a simple RC network - as I don't think you want the '555 triggering at the sound frequency rate. Or maybe you don't care for this application.
Flip R1 and the mic?
If we do this, I think the circuit makes a bit more sense. Q1 would then be OFF, and the TRI input to the '55 high in the absence of sound, since the base of Q1 would be pulled to ground by R1.
Sound would then turn on Q1, which would trigger the timer, creating a pulse that causes the 7474 to toggle. So long as the pulse duration is longer than the period of the sound wave, I don't think any additional filtering is needed.